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GeneE
26 sources retrieved · Most recent: April 2026 · Index updated 14 days ago
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CXCL9
C-X-C motif chemokine ligand 9
Chromosome 4 · 4q21.1
NCBI Gene: 4283Ensembl: ENSG00000138755.6HGNC: HGNC:7098UniProt: Q07325
229PubMed Papers
20Diseases
0Drugs
0Pathogenic Variants
FUNCTIONAL ROLE
Hub Gene
RESEARCH IMPACT
Trending
DATA QUALITY
✓ Experimental GO Evidence✓ Swiss-Prot Reviewed
adenylate cyclase-activating G protein-coupled receptor signaling pathwayregulation of cell population proliferationpositive regulation of release of sequestered calcium ion into cytosolprotein bindingneoplasmmelanomaVitiligobreast cancer
✦AI Summary

CXCL9 (C-X-C motif chemokine ligand 9) is a key immune-regulatory chemokine that primarily functions in T cell recruitment and activation through binding to the CXCR3 receptor 1. The protein is strongly induced by interferon-gamma (IFN-γ) and plays a central role in Th1-type immune responses, correlating with tissue infiltration of T cells 1. CXCL9 acts as a chemoattractant for activated T cells, particularly CXCR3+CD8+ T cells, and is involved in autoimmune processes such as vitiligo where it promotes melanocyte destruction through recruitment of cytotoxic T cells 2. In the tumor microenvironment, CXCL9 influences tumor-associated macrophage polarity and serves as a prognostic indicator, with its expression affecting treatment responses and patient outcomes 3. The chemokine demonstrates therapeutic potential in cancer immunotherapy, where CXCL9-engineered dendritic cells enhance T cell infiltration and activation, improving immune checkpoint blockade efficacy in non-small cell lung cancer 4. CXCL9 also serves as a diagnostic biomarker for acute interstitial nephritis, with significantly elevated urinary levels in affected patients 5. Additionally, CXCL9 plays roles in vascular remodeling, neuroinflammation, and formation of tertiary lymphoid structures that predict immunotherapy responses 67.

Sources cited
1
CXCL9 binds to CXCR3 receptor and is induced by IFN-γ in Th1-type responses
PMID: 29893515
2
CXCL9 recruits CXCR3+CD8+ T cells in vitiligo autoimmune response
PMID: 37937817
3
CXCL9 influences tumor-associated macrophage polarity and serves as prognostic indicator
PMID: 38217023
4
CXCL9-engineered dendritic cells enhance T cell infiltration and immune checkpoint blockade
PMID: 38518770
5
Urinary CXCL9 serves as diagnostic biomarker for acute interstitial nephritis
PMID: 37395276
6
CXCL9 inhibits collagen deposition via CXCR3 in vascular remodeling
PMID: 35763380
7
CXCL9-secreting endothelial venules recruit T cells into tertiary lymphoid structures
PMID: 40543508
Disease Associationsⓘ20
neoplasmOpen Targets
0.12Weak
melanomaOpen Targets
0.11Weak
VitiligoOpen Targets
0.10Weak
breast cancerOpen Targets
0.10Weak
hepatocellular carcinomaOpen Targets
0.10Suggestive
Townes-Brocks syndromeOpen Targets
0.10Suggestive
infectionOpen Targets
0.10Suggestive
rheumatoid arthritisOpen Targets
0.09Suggestive
triple-negative breast cancerOpen Targets
0.09Suggestive
pancreatic adenocarcinomaOpen Targets
0.09Suggestive
gastric cancerOpen Targets
0.09Suggestive
systemic lupus erythematosusOpen Targets
0.09Suggestive
tropical spastic paraparesisOpen Targets
0.09Suggestive
ovarian cancerOpen Targets
0.08Suggestive
systemic sclerodermaOpen Targets
0.08Suggestive
interstitial lung diseaseOpen Targets
0.08Suggestive
non-alcoholic fatty liver diseaseOpen Targets
0.08Suggestive
nasopharyngeal carcinomaOpen Targets
0.08Suggestive
prostate cancerOpen Targets
0.08Suggestive
lobular neoplasiaOpen Targets
0.08Suggestive
Pathogenic Variants
No pathogenic variants reported on ClinVar for this gene.
View on ClinVar ↗
Related Genes
IL1RNProtein interaction100%IRF1Protein interaction100%CCL8Protein interaction100%ACKR3Protein interaction100%CCR2Protein interaction100%CXCL17Protein interaction100%
Tissue Expression6 tissues
Lung
100%
Heart
75%
Liver
48%
Bone Marrow
32%
Ovary
3%
Brain
3%
Gene Interaction Network
Click a node to explore
CXCL9IL1RNIRF1CCL8ACKR3CCR2CXCL17
PROTEIN STRUCTURE
Preparing viewer…
AlphaFoldAI-predicted · UniProt Q07325
View on AlphaFold ↗
Constraintⓘ
LOEUFⓘ
1.36LoF Tolerant
pLIⓘ
0.02Tolerant
Observed/Expected LoF0.69 [0.38–1.36]
RankingsWhere CXCL9 stands among ~20K protein-coding genes
  • #1,760of 20,598
    Most Researched229 · top 10%
  • #14,191of 17,882
    Most Constrained (LOEUF)1.36
Genes detectedCXCL9
Sources retrieved26 papers
Response time—
📄 Sources
26▼
1
CXCL9, CXCL10, CXCL11, and their receptor (CXCR3) in neuroinflammation and neurodegeneration.
PMID: 29893515
Adv Clin Exp Med · 2018
1.00
2
The IFN-γ-CXCL9/CXCL10-CXCR3 axis in vitiligo: Pathological mechanism and treatment.
PMID: 37937817
Eur J Immunol · 2024
0.90
3
Perforin Generated by CD8
PMID: 40120774
Gastroenterology · 2025
0.80
4
CXCL9 and its Receptor CXCR3, an Important Link Between Inflammation and Cardiovascular Risks in RA Patients.
PMID: 37542661
Inflammation · 2023
0.72
5
Deciphering tumor microenvironment: CXCL9 and SPP1 as crucial determinants of tumor-associated macrophage polarity and prognostic indicators.
PMID: 38217023
Mol Cancer · 2024
0.70